Concerning the Mtuka Member, M. moyowamkia represents the first formally described tetrapod from this unit. The Mtuka Member is comparatively less known than the Namba Member with a fauna represented by remains of lungfish, turtles, crocodyliforms, theropod dinosaurs, and additional sauropod remains [44]. Although the two members are superficially similar in faunal composition at higher taxonomic rankings, M. moyowamkia is decisively distinct from both R. bisepultus and S. songwensis based on anatomical comparisons of currently known common elements. The cervical vertebrae of M. moyowamkia lack the autapomorphies described for R. bisepultus (e.g., carotid processes, accessory tubercles near the capituloparapophyseal suture, deep fossa along the ventral surface of the diapophysis; [11]). The caudal vertebrae of M. moyowamkia and R. bisepultus differ in several regards: (1) the anterior caudal vertebrae in R. bisepultus are strongly procoelous whereas the anterior caudal vertebra are weakly procoelous in M. moyowamkia; (2) R. bisepultus exhibits a tubercle near the prezygapophysis on the spinoprezygapophyseal lamina; (3) the neural spine in M. moyowamkia is comparatively narrower and directed dorsally rather than posterodorsally in R. bisepultus; (4) R. bisepultus lacks the dorsolateral expansion of the posterior face of the middle caudal centrum that is unique in M. moyowamkia; (5) the postzygapophysis in M. moyowamkia is oriented obliquely and offset from the neural spine; and (6) the distal caudal vertebra is transversely narrower in M. moyowamkia. Additionally, none of the numerous recovered dorsal ribs of M. moyowamkia and R. bisepultus exhibit the anterior and posterior flanges that are characteristic of S. songwensis. The M. moyowamkia humerus is more anteroposteriorly expanded than seen in S. songwensis and lacks the autapomorphies seen in R. bisepultus [12]. Otherwise the M. moyowamkia and S. songwensis humeri are incomplete in such a way to prevent more confident comparisons (e.g., the S. songwensis humerus does not preserve the deltopectoral crest but this feature is preserved in M. moyowamkia). Finally, since M. moyowamkia is recovered in a phylogenetically distant position from both S. songwensis and R. bisepultus, which are recovered in positions more closely related to titanosaurians of the Late Cretaceous, the phylogenetic placement is consistent with the Mtuka Member as a distinct and older unit from the Namba Member [44, 50].
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